Tenebrioninae are a large subfamily of the darkling beetles (Tenebrionidae), containing, among others, the flour beetles. Many of these robust and usually mid-sized beetles have elytra with some sort of corrugation on the upperside. They eat both fresh and decaying vegetation, including vegetable produce, and several are commercially important pests of flour and other cereal products. Tenebrioninae are eaten by large invertebrates and small vertebrates, such as birds, rodents, and lizards.